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Recommendations
Three letters of recommendation must be submitted from research mentors, faculty advisors and/or employers acquainted with the applicant’s work in the major area of study. Please use the recommender’s professional e-mail address. Do not use personal e-mail addresses through services such as AOL, Hotmail or Yahoo. Upon submission of the recommender’s name and e-mail address, an e-mail will be sent to the recommender containing a web link. The recommender will then be able to complete the necessary recommendation.

Letters of recommendation must be submitted through the online system no later than the application due date. Please note that you may submit your application before all letters have been received in our system, though we advise you to monitor the submission of letters of recommendation to ensure all letters are received by the deadline. If you are concerned your letters will not be submitted on time, please reach out to dbbsphaadmissions@email.wustl.edu.

Applicants are asked whether they wish to waive the opportunity to view their recommendation letters. Selecting Yes indicates that you will not be able to view the content of the recommendation letter in the future. Selecting No indicates that you may wish to view it in the future. Your decision will be shared with your recommenders and designated program so that they understand whether you waived the ability to see the content of the letter. Once you make a selection, you cannot change it, so consider the choices carefully. 

Please note recommendation letters are used to help reach decisions on admission and are not generally retained as part of the academic record of a student who matriculates at Washington University in St. Louis.

Applicants may review the status of required supplemental materials, such as letters of recommendation by signing into the Status Portal of the online application. If a letter(s) have not been submitted, please contact the recommender(s) and ask them to complete by the application deadline. If your recommenders have not received the recommendation email request, you may resend the request via the Status Portal.
 
PhD Application Instructions
5
Yes
  
Test Scores

GRE:
The Graduate Record Examination (GRE) General test is no longer required by DBBS Programs, however it is optional for some. Please refer to the individual program web page (http://www.dbbs.wustl.edu/divprograms/Pages/Division-Programs.aspx) to see if GRE scores may be optionally submitted.  If so, scores must be from tests taken in the last five years.

Official GRE Scores: Washington University in St. Louis - School code 6929.​

English Proficiency Test Requirement for International Applicants:
As part of the application review, we are required to verify that each nonnative English speaking applicant has sufficient English language proficiency to succeed in the academic program.

The English proficiency test requirement is waived for citizens of Australia, Cameroon, Commonwealth Caribbean nations, Ghana, India, Ireland, Kenya, Liberia, New Zealand, Nigeria, Singapore, Uganda, the United Kingdom, Zambia, and Zimbabwe. It may also be waived for students who have completed three or more years of study in an English-medium academic program in any of the above countries, Canada, Hong Kong, South Africa, or the United States.

If you are required to provide proof of English proficiency, the application will prompt you to self-report English proficiency scores that are less than two years old. Official scores should be sent to Washington University in St. Louis at the time of application. We accept the following English proficiency tests:

English Proficiency Tests for Application to 2023-2024

Official TOEFL IBT -- Minimum Score 100: Send electronically through ETS to school code 6929

Official IELTS  --  Minimum Score 7.5: Send electronically by selecting Washington University in St. Louis Graduate Admissions
​​
Official Duolingo  --  Minimum Score 125: Send link with scores to dbbsphdadmissions@email.wustl.edu


PhD Application Instructions
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GPA
Calculate the GPA for each school attended on a 4.0 scale.  To convert:

GPA/max. score X 4 = reported score to us (rounded to the nearest 0.1)

For example:
          On a 100 point scale: 92/100 = .92 X 4 = 3.7
          On a 5 point scale: 3.9/5 = 0.78 X 4 = 3.1

PhD Application Instructions
7
No
  
Transcripts
  • Prior to submitting your application, you are required to upload unofficial transcript(s) in PDF format from each insitution attended. Transcript must have your name, the institution name, and be translated to English when necessary. 
  • Scan the transcript at high resolution and verify each page can be read before uploading it.  Upload the transcript from each institution as a single document. Each individual university transcript must be uploaded as one file and not page by page.
  • Official transcripts are not required for the review process. If you are offered admission and accept the offer you are then required to submit an official final transcript noting degree confirmation from each degree granting institution you have attended.
  • DBBS reserves the right to rescind any offer of admission if discrepancies are found between your uploaded transcript and the official transcript.
PhD Application Instructions
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Essays
Applicants are encouraged to give considerable time and thought to writing the two essays. (Some schools ask for a general “personal statement”, however DBBS requires that the applicant respond to specific essay prompts.) The Admissions Committee evaluates a number of characteristics in order to assess a candidate’s ability to benefit from and contribute to their selected program. These include academic ability, research experience, leadership, teamwork, communication skills and other personal qualities. The essays are an opportunity to help the Committee relate to the applicant’s distinctive values, motivation, and perspectives and obtain as much information as possible about their previous research experience.
 
Essay 1: a) describe your research project(s) and specific contributions to the work; b) discuss your motivations for graduate study; and c) relate a fundamental biological, biomedical, or data science question that most intrigues you, highlighting potential Washington University faculty mentors.
 
Essay 2: drawing on your past experiences within or beyond your research, please provide a statement showcasing your strengths as an applicant and articulating why you will be an outstanding graduate student.​
 
Applicants are requested to upload each essay as either a PDF or Word document. Applicants are highly encouraged to submit no more than two (2) pages per essay (or approximately 8000 characters including spaces). Please address each essay question separately. Do not insert a personal statement, CV, resume or publication in lieu of the two essay questions.
PhD Application Instructions
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Prerequisites
DBBS welcomes applicants with outstanding baccalaureate training in natural, physical or engineering sciences that are committed to a career in research. The strongest applicants will have had significant research experience and have a strong undergraduate record not only in their area of concentration, but also in related disciplines. For example, some of our programs will give preference to applicants who have strong training not only in biology, but also 3 to 6 semesters of chemistry (including biochemistry) as well as appropriate course work in calculus and physics (1 to 2 semesters each). Other programs, depending on their emphasis, will consider individuals with less extensive backgrounds in chemistry, physics, psychology or mathematics but with related training in biology. For example, Biochemistry, Biophysics, and Structural Biology gives preference to applicants that have completed one semester of physical chemistry or an equivalent course.

Minimum Requirements:
  • All applicants must have a U.S. bachelor’s degree or its equivalent from an accredited institution.
  • The Graduate Record Examination (GRE) General test is not required to apply to DBBS.  Please refer to "Test Scores" below for additional information.
  • Proficiency in English is required of all applicants. Applicants whose native language is not English must demonstrate English proficiency and are required to provide an official ELP exam score report. Please refer to "Test Scores" below for additional information.​  
PhD Application Instructions
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Application
All applicants must have a U.S. bachelor’s degree or its equivalent from an accredited institution.

The application consists of:
 
  • Demographics
  • Three letters of recommendation
  • Academic history
  • Test Scores 
    • GRE (not required, but optional for some programs)
    • TOEFL, IELTS, or Duolingo (if applicable)
  • Two essays 
  • 10 word  undergrad research description & 10 word descripotion of graduate research interest
  • Listing of Major coursework
  • Work and Research Experience
  • Awards/Honors
  • Publication list (if any)

Applicants who have applied in the previous year must submit a new application and answer Yes to the question, "Have you previously applied to a DBBS program​ at Washington University?".

All application materials sent to us by the applicant become the property of the Roy and Diana Vagelos Division of Biology and Biological Sciences at Washington University and cannot be returned to the applicant or forwarded to another school/department.

PhD Application Instructions
1
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Deadlines

The application deadline for Fall 2022 enrollment is December 1, 2021 @ 11:59 PM PST. All applicants are encouraged to submit ealry.  Submission of the application prior to November 1, 2021 will allow for a discounted application fee.

PhD Application Instructions
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Mandatory Clinical Mentorship Orientation

Date/Time

Location

Speaker

Title

Tues, Nov 22, ​2022

2:00 pm – 3:00 pm*

FLTC 205

Ryan Fields, MD

Professor of Surgery

Mandatory

Clinical Mentorship Orientation

Pathobiology of Human Disease States Course
4
Yes
  
Past Course Modules
Year Modules
1992 Sickle Cell Anemia
Malaria
Diabetes
1993 Cystic Fibrosis
Coronary Artery Disease
AIDS
1994 Sickle Cell Anemia
Emphysema
Acute Leukemia
1995 Multiple Endocrine Neoplasia
Undue Susceptibility to Infection
Alzheimer’s Disease
1996 Sickle Cell Disease
Multiple Sclerosis
Epstein Barr Virus
1997 Osteoporosis
Thrombophilia
Congenital Malformations
1998 Sickle Cell Disease
Hepatitis C
Chronic Myelogenous Leukemia
1999 Rheumatoid Arthritis
AIDS
Prostate Cancer
2000 Sickle Cell Disease
Epstein Barr Virus
Heart Failure
2001 Alzheimer’s Disease
Chronic Myelogenous Leukemia
SLE (Lupus)
2002 Heart Failure
Diabetes
Mellitus Preeclampsia
2003 SLE/Rheumatoid Arthritis
Chronic Myelogenous Leukemia
Asthma
2004 AIDS
Heart Failure
Preeclampsia
2005 Alzheimer’s Disease
Breast Cancer
Rheumatoid Arthritis
2006 Sickle Cell Anemia
Congestion Heart Failure
High Risk Pregnancy
2007 Rheumatoid Arthritis
Chronic Myelogenous Leukemia
Alzheimer’s Disease
2008 Cardio Disease/Heart Failure
Diabetes & Metabolism
Osteoporosis/Other Bone Disorders
2009 Sickle Cell Anemia
AIDS
Major Depressive Disorders
2010 Alzheimer’s Disease
Cardio Disease/Heart Failure
Diabetes
2011 Chronic Myelogenous Leukemia
Cystic Fibrosis
High Risk Pregnancy
2012 Rheumatoid Arthritis
Cardiovascular Disease
Malaria
2013 Diabetes
High Risk Pregnancy
Depression
2014 Translating Global Health
Cardiovascular Disease
Chronic Myelogenous Leukemia
2015 Immunotherapy
High Risk Pregnancy
Alzheimer’s Disease
2016 Evidence Based Medicine/Sickle Cell Disease
Diabetes
Cardiovascular Disease
2017 Immunotherapy
Alzheimer’s Disease
Hearing Loss
2018 High Risk Pregnancy
Pain Management
Cardiovascular Disease
2019 Immunotherapy
Diabetes
Retina
2020 Alzheimer’s Disease
Hearing Loss
Cardiovascular Disease
2021 Sickle Cell Disease
High Risk Pregnancy
Pain Management
Pathobiology of Human Disease States Course
5
Yes
  
Module 3 - Immunotherapy

Module Leaders: 

Nathan Singh, MD, MS   Assistant Professor of Medicine, Division of Oncology
Todd Fehniger, MD, PhD   Professor of Medicine, Division of Oncology

Date/Time

Location

Speaker

Title

Patient Visitor (Y/N)

Tues, Oct 25, 2022

1:30 – 3:00 pm

FLTC 205

​Nathan Singh, MD

Assistant Professor,  

Department of Medicine,

Division of Oncology

 

Title

 

Y

Thurs, Oct 27, 2022

1:30 – 3:00 pm

FLTC 205

Carl DeSelm, MD, PhD

Assistant Professor,

Department of Radiation Oncology

 

Title

 

 

Tues, Nov 1, 2022

1:30 – 3:00 pm

FLTC 205

Todd Fehniger, MD, PhD

Professor,

Department of Medicine,

Division of Oncology

 

Title

 

Y

Thurs, Nov 3, 2022

1:30 – 3:00 pm

FLTC 205

Greg Wu, MD, PhD

Associate Professor,

Department of Neurology

 

Title

 

Y

Tues, Nov 8, 2022

1:30 – 3:00 pm

FLTC 205

Jeff Ward, MD, PhD

Assistant Professor,  

Department of Medicine,

Division of Oncology

 

Title

 

Y

Thurs, Nov 10, 2022

1:30 – 3:00 pm

FLTC 205

Amanda Cashen, MD

Professor,

Department of Medicine,

Division of Oncology

 

Title

 

Y

Tues, Nov 15, 2022

1:30 – 3:00 pm

TBD*

Field Trip!

 

Title

 

 

Thurs, Nov 17, 2022

(Wrap-up)

1:30 – 3:00 pm

FLTC 205

All Speakers (if available)

Faculty Overview and course discussion -        Lunch served

 

Pathobiology of Human Disease States Course
3
Yes
  
Module 2 - Kidney Disease

Module Leader: Carmen Halabi, MD, PhD Assistant Professor of Pediatrics, Division of Nephrology

Date/Time

Location

Speak​er

Title

Patient Visitor (Y/N)

Tues, Sep 27, 2022

1:30 – 3:00 pm

FLTC 205

Field Trip

Dialysis Center*

Carmen Halabi, MD, PhD

Assistant Professor of Pediatrics, Division of Nephrology

 

Introduction to kidney disease/tour of the pediatric dialysis unit

 

Y

Thurs, Sep 29, 2022

1:30 – 3:00 pm

FLTC 205

Michael Rauchman, MDCM

Chromalloy Professor of Medicine,

Division of Nephrology

 

Genetic basis of congenital anomalies of the kidney and urinary tract

 

 

Tues, Oct 4, 2022

1:30 – 3:00 pm

FLTC 205

Andrew Malone, MB, BCh

Assistant Professor of Medicine, Transplant Nephrology,

Division of Nephrology

 

Kidney disease progression

 

 

Thurs, Oct 6, 2022

1:30 – 3:00 pm

FLTC 205

Parker Wilson, MD, PhD

Assistant Professor of Pathology & Immunology

 

Multi-omic analyses of human diabetic kidney disease

 

 

Tues, Oct 11, 2022

No Class – Fall Break

n/a

n/a

 

n/a

 

n/a

Thurs, Oct 13, 2022

1:30 – 3:00 pm

FLTC 205

Leslie Gewin, MD

Associate Professor of Medicine,

Division of Nephrology

Application of single cell technologies in kidney transplantation

 

 

Tues, Oct 18, 2022

1:30 – 3:00 pm

FLTC 205

Jeff Miner, PhD, FASN

Eduardo and Judith Slatopolsky Endowed Professor of Medicine,

Division of Nephrology

 

Novel therapies for Alport’s syndrome

 

 

Thurs, Oct 20, 2022

(Wrap-up)

1:30 – 3:00 pm

FLTC 205

All Speakers (if available)

Faculty Overview and course discussion -        Lunch served

 

Pathobiology of Human Disease States Course
2
Yes
  
Module 1 - Diabetes

Module Leader:  Tom Baranski, MD, PhD Professor of Medicine and Developmental Biology, Division of Endocrinology

Date/Time

 

Location

Speaker

Title

Patient Visitor (Y/N)

Tues, Aug 30, 2022

1:30 – 3:00 pm

FLTC 205

Colin Nichols, PhD

Carl Cori Professor, Department of Cell Biology and Physiology

Director, Center for the Investigation of Membrane Excitability Diseases

Monogenic Diabetes

 

Thurs, Sep 1, 2022

1:30 – 3:00 pm

FLTC 205

Kevin Bauerle, MD, PhD

Instructor in Medicine

Division of Endocrinology

Obesity and Insulin Resistance

 

Tues, Sep 6, 2022

1:30 – 3:00 pm

FLTC 205

Paul Hruz, MD, PhD

Associate Professor of Pediatrics

Divisions of Cell Biology & Physiology and Endocrinology and Diabetes

Insulin Action-lessons from Transgenic Animals

 

Thurs, Sep 8, 2022

1:30 – 3:00 pm

FLTC 205

Tom Baranski, MD, PhD

Professor of Medicine and Developmental Biology

Division of Endocrinology

Complications of Diabetes

Y

Tues, Sep 13, 2022

1:30 – 3:00 pm

FLTC 205

Fumi Urano, MD, PhD

Samuel E. Schechter Professor of Medicine, Professor of Medicine and of Pathology and Immunology

Division of Endocrinology

Wolfram Syndrome

Y

Thurs, Sep 15, 2022

1:30 – 3:00 pm

FLTC 205

Janet McGill, MD, MA, FACE

Professor of Medicine

Division of Endocrinology

Introduction to Clinical Diabetes

Y

Tues, Sep 20, 2022

1:30 – 3:00 pm

FLTC 205

Jeff Millman, PhD

Associate Professor of Medicine and Biomedical Engineering

Division of Endocrinology

Pluripotent Technologies and Beta Cells

 

Thurs, Sep 22, 2022

(Wrap-up)

1:30 – 3:00 pm

FLTC 205

All Speakers (if available)

Faculty Overview and course discussion -        Lunch served

 

Pathobiology of Human Disease States Course
1
Yes
  
William H. Danforth Fellowship in Plant Sciences

Natasha Bilkey - Plant and Microbial Biosciences

Ryan Calcutt - Plant and Microbial Biosciences

Jennette Codjoe - Plant and Microbial Biosciences

Kristen Edgeworth - Plant and Microbial Biosciences

Kiona Elliott - Plant and Microbial Biosciences

Wen-Hsi KuoEvolution, Ecology and Population Biology

Erin Mattoon​ - Plant and Microbial Biosciences

William McHarguePlant and Microbial Biosciences

Kari M​illerPlant and Microbial Biosciences

Lily O'ConnorPlant and Microbial Biosciences

External Fellowship Awardees
38
Yes
  
National Science Foundation

Maliayah Adkins-Threats​Developmental, Regenerative & Stem Cell Biology

Mahal Bugay - Evolution, Ecology and Population Biology

Mandy Chan - Immunology

Jennette Codjoe - Plant and Microbial Biosciences

Melissa Cook - Immunology

Emily Coonrod - Human & Statistical Genetics

Sophia DeGeorgia - Molecular Cell Biology

Kiona Elliott - Plant and Microbial Biosciences

Paula Godoy - Computational & Systems Biology

Daniel Griffith - Computational & Systems Biology

Gregory Harriso​n​ - Molecular Microbiology and Microbial Pathogenesis

Christy Hoffman - Human & Statistical Genetics

Sarah Hurt - Molecular Genetics & Genomics

Lauren Johnson - Evolution, Ecology and Population Biology

Elizabeth Kennedy - Molecular Microbiology and Microbial Pathogenesis

Abigail Ki​mball Molecular Microbiology and Microbial Pathogenesis

Jennifer Lawrence - Neurosciences

Kari Miller - Plant and Microbial Biosciences

Kayla Nygaard - Molecular Genetics and Genomics

Sarah PyeMolecular Microbiology and Microbial Pathogenesis

Ananda RankinMolecular Microbiology and Microbial Pathogenesis

India Reiss - Neurosciences

Nicole Rivera-EspinalMolecular Microbiology and Microbial Pathogenesis

Celine St. Pierre - Human & Statistical Genetics

Michael Strickland - Neurosciences

Chayla Vazquez - Neurosciences

Daniel Veronese Paniagua​ Developmental, Regenerative & Stem Cell Biology

Hung Vuong​ - Molecular Microbiology and Microbial Pathogenesis

External Fellowship Awardees
24
Yes
  
National Institute of Health

Shashank Anand - Biomedical Engineering

John Baer​ - Molecular Cell Biology

Keenan​ Bates Developmental, Regenerative & Stem Cell Biology

Zach BellerComputational & Systems Biology

Druv Bhagavan - Biomedical Engineering

Rebecca Brenner - Neurosciences

Lindsey Brier - Neurosciences

Cat Camacho - Neurosciences

Kitra Cates - Molecular Genetics & Genomics

Robert Chen​ - Computational & Systems Biology

Christopher Chermside-Scabbo - Computational & Systems Biology

Samantha Chin - Molecular Cell Biology

Diana Christian - Neurosciences

Kolin Clark - Molecular Genetics & Genomics

Matthew Cruz - Biochemistry, Biophysics, and Structural Biology

Celia Cubitt​ - Immunology

Peppar Cyr - Neurosciences

Anna Damato​ - Neurosciences

Luke Diorio-Toth - Computational & Systems Biology

Jared Ele​nbaas​ - Molecular Genetics & Genomics

Yan-Yang Feng - Biomedical Engineering

Ryan FriedmanComputational & Systems Biology

Steven GrigsbyMolecular Microbiology & Microbial Pathogenesis

Ariel Hernandez-Leyva - Computational & Systems Biology

Graham Hogg - Immunology

Alexandra Houston-Ludlam​ - Human and Statistical Genetics

Po wei 'Billy' Kang​ - Biomedical Engineering

Marlene Kanmogne - Neurosciences

Clara Kao - Molecular Genetics & Genomics

Elizabeth KennedyMolecular Microbiology & Microbial Pathogenesis

Joseph Krambs - Molecular Genetics & Genomics

Peter Lambert - Neurosciences

Varintra Lander - Immunology

Jamison LeidDevelopmental, Regenerative & Stem Cell Biology

Nicolas Ledru - Molecular Genetics & Genomics

Aisha Lee - Immunology

Joseph Lin - Neurosciences

Liza Loza​ Molecular Microbiology & Microbial Pathogenesis​

Manasi Malik - Molecular Cell Biology

Sindhu Manivasagam​ - Neurosciences

Megan MichieBiochemistry, Biophysics, and Structural Biology

James "Russell" Moore - Neurosciences

Bernard Mulvey​ - Neurosciences

Makenzie Norris - Neurosciences

Takihiro Ohara - Immunology

Vincent Peng - Immunology

India Reiss - Neurosciences

James Shepherdson - Computational & Systems Biology

Sarah Speck - Molecular Cell Biology

Jennifer Tran - Immunology

Anna Trier - Immunology

Brittany Townley - Molecular Cell Biology

Ryan Valdez - Plant & Microbial Biosciences

Jace Webster​ - Human & Statistical Genetics

Emma Winkler - Immunology

Renee Wu -Immunology

Christopher Yoon - Molecular Genetics & Genomics

Yiming ZhangDevelopmental, Regenerative & Stem Cell Biology

Cyrus Zhou - Molecular Genetics & Genomics

External Fellowship Awardees
20
Yes
  
American Heart Association

Junedh Amrute - Computational and Systems Biology

Kendall Burks - Molecular Genetics and Genomics

Jiwo​n Yi​​ - Neurosciences

Alex Legaria Macal - Neurosciences

External Fellowship Awardees
2
Yes
  
Gary K. Ackers Fellowship

Drake Jensen - ​Computational and Molecular Biophysics

Brittan​y Smith​ - Computational and Molecular Biophysics

External Fellowship Awardees
12
Yes
  
Childrens Tumor Foundation

​Michelle W​egscheid​ - Neurosciences

External Fellowship Awardees
5
Yes
  
National Aeronautics and Space Administration (NASA)

Cameron​ Jay Sargent - Biochemistry

External Fellowship Awardees
20
Yes
  
Agency for Science, Technology, and Research
Ying Shi​ang Lim​ - Immunology
External Fellowship Awardees
1
Yes
  
Schneiderman Fellowship

Ross McKi​nney​ - Neurosciences​

External Fellowship Awardees
30
No
  
Canadian Institutes of Health Research

Jad​ Belle​Molecular Genetics and Genomics

External Fellowship Awardees
4
Yes
  
Kwanjeong Educational Foundation

​Seunghyun (S​hawn) Lee​ - Immunology

External Fellowship Awardees
14
Yes
  
Male Contraceptive Inititiative (MCI)

​Max ​Lyon​ - Molecular Cell Biology

External Fellowship Awardees
16
Yes
  
MilliporeSigma Predoctoral Fellowship in Honor of Dr. Gerty T. Cori

​Nicole Fazio - Computational and Molecular Biophysics

Catherine Knoverek - Biochemistry, Biophysics, and Structural Biology

Sukrit ​Singh - Computational and Molecular Biophysics

External Fellowship Awardees
17
Yes
  
Molecular Science Software Institute

Matthew Ward - Molecular Cell Biology

​Micha​el Ward​ - Molecular Cell Biology

External Fellowship Awardees
18
Yes
  
Monsanto Excellence Fund for Graduate Fellowships in Life Sciences

Gervette Penny - Molecular Genetics & Genomics

Patric​ia Walker - Plant & Microbial Biosciences

External Fellowship Awardees
19
Yes
  
Spencer T. and Ann W. Olin Fellowship for Women in Graduate Study

Emma Frawley - Evolution, Ecology and Population Biology

Alexandra​ Scott​ - Molecular Genetics and Genomics

External Fellowship Awardees
33
Yes
  
Chancellor Fellows

Paula Godoy - Computational and Systems Biology

Taylor Harris​ - Plant and Microbial Biosciences

Dymonn Johnson​ - Molecular Genetics and Genomics

Joseph Kr​ambs - Molecular Genetics and Genomics

External Fellowship Awardees
10
Yes
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